Dryopteris hybrid complex

The fern genus Dryopteris is known to be one of the most freely-hybridizing of fern genera.

Parent species

• Dryopteris expansa
• Dryopteris goldiana
• Dryopteris intermedia
• Dryopteris ludoviciana
• Dryopteris marginalis

Hybrid species

• Dryopteris carthusiana (D. intermedia × unknown parent; allotetraploid)
• Dryopteris campyloptera (D. intermedia × D. expansa; allotetraploid)
• Dryopteris celsa (D. goldiana × D. ludoviciana; allotetraploid)
• Dryopteris clintoniana (D. cristata × D. goldiana; allohexaploid)
• Dryopteris cristata (D. goldiana × unknown parent; allotetraploid)

Other hybrids

• Dryopteris × bootii (D. cristata × D. intermedia; triploid)
• Dryopteris × critica (D. borreri × D. filix-mas)^[1]
• Dryopteris × complexa aggregate (D. filix-mas and D. affinis; tetraploid)^[1]
• Dryopteris × convoluta (D. cambrensis × D. filix-mas)^[1]
• Dryopteris × deweveri (D. dilatata × D. carthusiana)^[1]
• Dryopteris × neo-wherryi (D. goldiana × D. marginalis; diploid)
• Dryopteris × triploidea (D. carthusiana × D. intermedia; triploid)

References

1. Murphy, Rosaline J.; Page, Christopher N.; Parslow, Rosemary E.; Bennallick, Ian J. (2012). Ferns, Clubmosses, Quillworts and Horsetails of Cornwall and the Isles of Scilly. Truro: ERCCIS. ISBN 978-1-902864-07-5.