Hemipenthes

Hemipenthes
Hemipenthes maurus
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Diptera
Suborder:	Brachycera
Infraorder:	Asilomorpha
Superfamily:	Asiloidea
Family:	Bombyliidae
Subfamily:	Anthracinae
Tribe:	Exoprosopini
Genus:	Hemipenthes Loew, 1861
Type species
Musca morio Linnaeus, 1758
Synonyms
Isopenthes Osten Sacken, 1877

Hemipenthes is a large genus of flies belonging to the family Bombyliidae (bee-flies). There are many described species, distributed throughout the Holarctic and Palearctic Ecozones. These are small to large robust flies with a body length of 5–14 mm. They can be distinguished from similar genera (Villa) by their wing venation.^[1]

Larvae are hyperparasites on parasitic Hymenoptera.^[1]

Species

Includes fossil species.

• H. bigradatus (Loew, 1869) - Nearctic: USA (California, New Mexico). Neotropical: Cuba, Bahamas, Jamaica, Puerto Rico.
• H. blanchardiana (Jaennicke, 1867) - Nearctic: Mexico (Distrito Federal, Guanajuato, Guerrero, México, Puebla, Sonora), USA (Arizona, California, Texas).
• H. castanipes Bigot, 1892 - Nearctic:
• H. catulina (Coquillett, 1894) - Nearctic: USA (California, Colorado, Idaho, Illinois, Indiana, Iowa, Michigan, Minnesota, Montana, New Jersey, New Mexico, New York, Ohio, Pennsylvania, Utah, Washington, Wisconsin, Wyoming).
• H. celeris (Wiedemann, 1828) - Nearctic: USA (Kentucky)
• H. chimaera (Osten Sacken, 1887) - Nearctic: Mexico (Guerrero, Sonora), USA (Arizona)
• H. comanche (Painter in Painter & Painter, 1962) - Nearctic: USA (Arizona, California, Colorado, Nevada, New Mexico, Utah).
• H. constituta Walker, 1852 - Neotropical: South America.
• H. curta (Loew, 1869) - Nearctic: Mexico (Colima, Guerrero, Jalisco, Morelos, Nuevo León, Sinaloa, Veracruz-Llave), USA (Arizona, California, New Mexico, Texas). Neotropical: Mexico (Oaxaca, Yucatán).
• H. differens (Hall, 1976) - Neotropical: Chile.
• H. discolor (Hall, 1976) - Neotropical: Chile.
• H. ditaenia (Wiedemann, 1828) - Neotropical: Argentina, Chile, Peru, Uruguay.
• H. divisa (Walker) - Neotropical: South America.
• H. edwardsii (Coquillett, 1894) - Nearctic: Canada (British Columbia), USA (California).
• H. edwarsi (Oldroyd, 1938) - Neotropical: Chile.
• H. epilais (Wiedemann, 1828) - Neotropical: Brazil
• H. ethiops (Greathead, 1967) - Afrotropical: Ethiopia
• H. eumenes (Osten Sacken, 1887) - Nearctic: Canada (Alberta), Mexico (Sonora), USA (Arizona, California, Colorado, Idaho, Montana, Nevada, New Mexico, Oregon, Washington). Neotropical: Puerto Rico.
• H. eversmanni Zaitzev and Madra Mandicencio, 1966 - Palaearctic: Kazakhstan, Moldova, Russia, Ukraine.
• H. exoprosopoides Paramonov, 1928 - Palaearctic: Armenia, Azerbaijan, China (Sichuan), Iran, Kyrgyz Republic, Tajikistan, Turkmenistan, Uzbekistan.
• H. extensa Wulp, 1888 - Neotropical: Argentina.
• H. floridanus (Macquart, 1850) - Nearctic: USA (Florida, Georgia).
• H. gabbroensis (Handlirsch, 1907) - Palaearctic: Italy (Miocene).
• H. galapagensis (Painter & Painter, 1974) - Neotropical: Argentina.
• H. galathea (Osten Sacken, 1886) - Neotropical: Costa Rica
• H. gaudanica Paramonov, 1927 - Palaearctic: China (Xinjiang), Iran, Kyrgyz Republic, Mongolia, Tajikistan, Turkmenistan, Uzbekistan.
• H. gayi (Macquart, 1840) - Neotropical: Chile.
• H. gussakovskyi Zaitzev, 1966 - Palaearctic: Tajikistan, Turkmenistan
• H. hamifera caucasica Zaitzev, 1966 - Palaearctic: Armenia, Azerbaijan, Gruzia, Iran.
• H. hamifera (Loew, 1854) - Palaearctic: Armenia, Azerbaijan, Bulgaria, China (Jiangsu, Nei Monggol, Xinjiang), France, Greece, Gruzia, Iran, Italy (incl. Sicily), Kazakhstan, Kyrgyz Republic, Mongolia, Russia (ES, WS), Spain, Tajikistan, Turkey, Turkmenistan, Uzbekistan, Yugoslavia.
• H. ignea (Macquart, 1846) - Neotropical: Colombia, Guyana, Panama.
• H. incisiva (Walker, 1852) - Nearctic: Mexico (Guerrero)
• H. inops (Coquillett, 1887) - Nearctic: USA (Arizona, California, Colorado, Oregon, Utah, Washington).
• H. jaennickeana (Osten Sacken, 1886) - Nearctic: Mexico (Morelos, Sonora), USA (Arizona, California, Colorado, Idaho, Montana, Nevada, New Mexico, Oregon, Texas, Utah).
• H. jezoensis (Matsumura, 1916) - Oriental: Taiwan. Palaearctic: Japan.
• H. lepidota (Osten Sacken, 1887) - Nearctic: Canada (Alberta), Mexico (Baja California Norte, Baja California Sur, Chihuahua, Guerrero, Puebla, San Luis Potosí, Sonora, Tamaulipas), USA (Arizona, California, Colorado, Idaho, Louisiana, Nevada).
• H. leucocephala (Wulp, 1882) - Neotropical: Argentina.
• H. martinorum (Painter in Painter & Painter, 1962) - Nearctic: Mexico (Michoácan de Ocampo)
• H. maurus (Linnaeus, 1758) - Palaearctic
• H. melaleuca (Wiedemann, 1828) - Neotropical: Argentina, Uruguay.
• H. melana Bowden, 1965 - Oriental: Nepal.
• H. mesasiatica Zaitzev, 1962 - Palaearctic: Tajikistan, Turkmenistan, Uzbekistan.
• H. micromelas (Bigot, 1892) - Neotropical: Argentina
• H. minas (Macquart, 1850) - Neotropical: Argentina, Brazil (Minas Gerais).
• H. mischanensis Paramonov, 1927 - Palaearctic: Armenia.
• H. montanorum (Austen, 1936) - Oriental: China (Yunnan). Palaearctic: China (Qinghai, Sichuan, Xizang).
• H. morio (Linnaeus, 1758) - Nearctic, Oriental, Palaearctic.
• H. nitidofasciata (Portschinsky, 1892) - Palaearctic: Kyrgyz Republic, Russia, Tajikistan.
• H. noscibilis (Austen, 1936) - Palaearctic: China (Xizang).
• H. nudiuscula (Thomson, 1869) - Neotropical: Panama.
• H. pamirensis Zaitzev, 1962 - Palaearctic: Mongolia, Tajikistan, Turkmenistan, Uzbekistan.
• H. panfilovi Zaitzev, 1981 - Palaearctic: Russia
• H. pauper (Becker, 1916) - Palaearctic: Algeria, Egypt.
• H. pima (Painter, 1962) - Nearctic: Mexico (Sinaloa, Sonora), USA (Arizona, New Mexico, Texas).
• H. pleuralis (Williston, 1901) - Nearctic: Mexico (Colima, Guerrero, Morelos, Nayarit, Puebla). Neotropical: Mexico (Chiapas).
• H. praecisa (Loew, 1869) - Palaearctic: China (Beijing, Hebei, Nei Monggol), Kyrgyz Republic, Mongolia, Russia, Tajikistan, Turkmenistan, Uzbekistan.
• H. provincialis (Handlirsch, 1907) - Palaearctic: France (Oligocene)
• H. pullata (Coquillett, 1894) - Nearctic: USA (Arizona, California).
• H. referens (Walker, 1852) - Oriental: India, Myanmar, Pakistan.
• H. robusta Zaitzev, 1966 - Palaearctic: Armenia, Azerbaijan, China, Gruzia, Kazakhstan, Kyrgyz Republic, Tajikistan, Turkmenistan, Uzbekistan.
• H. ruficollis (Bigot, 1892) - Neotropical: Argentina, Venezuela.
• H. sagata Loew - Nearctic: Mexico, USA (Arizona, Mississippi, Missouri, Texas). Neotropical: Costa Rica, Honduras, Panama.
• H. scylla (Osten Sacken, 1887) - Nearctic: Mexico (Guanajuato, Sonora), USA (Arizona, Texas). Neotropical: Venezuela.
• H. semifucata (Hall, 1976) - Nearctic: USA (Arizona).
• H. seminigra Loew, 1869 - Nearctic: Canada (Saskatchewan), USA (Utah)
• H. sinuosa (Wiedemann, 1821) - Nearctic: USA
• H. splendida Zaitzev, 1962 - Palaearctic: Kazakhstan, Kyrgyz Republic, Tajikistan, Turkmenistan.
• H. subarcuata Loew, 1871 - Palaearctic: Tajikistan, Tunisia.
• H. subvelutina Loew, 1871 - Palaearctic: Armenia, Azerbaijan, China (Shandong), Gruzia, Iran, Mongolia, Kazakhstan, Kyrgyz Republic, Tajikistan, Turkey, Turkmenistan, Uzbekistan.
• H. tarapacensis (Hall, 1976) - Neotropical: Chile.
• H. tenuirostris (Macquart, 1850) - Neotropical: Chile, Peru.
• H. tertiaria (Handlirsch, 1907) - Palaearctic: Germany (Oligocene).
• H. tushetica Zaitzev, 1966 - Palaearctic: Armenia, Azerbaijan, China (Nei Monggol, Qinghai, Xinjiang), Gruzia, Iran.
• H. tusheticus mongolicus Zaitzev, 1980 - Palaearctic: Mongolia.
• H. velutina (Meigen, 1820) - Oriental: Pakistan. Palaearctic: Albania, Algeria, Armenia, Austria, Azerbaijan, Belgium, Bosnia-Hercegovina, Bulgaria, China (Jiangsu, Nei Monggol, Qinghai, Xinjiang), Croatia, Cyprus, Czech Republic, Egypt, France, Germany, Greece, Hungary, Iran, Israel, Italy, Lebanon, Macedonia, Moldova, Mongolia, Morocco, Poland, Portugal, Romania, Russia (SET), Slovakia, Slovenia, Spain (incl. Mallorca), Switzerland, Syria, Tunisia, Turkey, Turkmenistan, Ukraine, Yugoslavia.
• H. villeneuvi François, 1970 - Palaearctic: France, Greece, Italy.
• H. vockerothi François, 1969 - Palaearctic: Spain.
• H. webberi (Johnson, 1919) - Nearctic: Canada (Ontario, Quebec), USA (Connecticut, Kentucky, Massachusetts, Vermont, Wyoming).
• H. wilcoxi (Painter, 1933) - Nearctic: USA (California, Washington).
• H. yaqui (Painter in Painter & Painter, 1962) - Nearctic: Mexico (Sonora), USA (Arizona)

^[2][3]

References

1. Hull, F.M. (1973). Bee flies of the world. The genera of the family Bombyliidae. Washington: Smithsonian Institution Press. pp. 687 pp. ISBN 0-87474-131-9.
2. "World Catalog Family Bombyliidae Latreille (part 1 (Oligodraninae through Bombyliinae, p. 1-206)" (PDF). Bishop Museum. 1999. pp. 1–206. Retrieved 12 January 2009.
3. "World Catalog Family Bombyliidae Latreille (part 1 (Oligodraninae through Bombyliinae, p. 1-206) - Revised Edition" (PDF). Bishop Museum. September 2003. pp. 1–206. Retrieved 12 January 2009.