Lamiales
Lamiales | |
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Galeopsis speciosa | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Angiosperms |
Clade: | Eudicots |
Clade: | Asterids |
Clade: | Lamiids |
Order: | Lamiales Bromhead[1] |
Families[2] | |
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The Lamiales are an order in the asterid group of dicotyledonous flowering plants. It includes about 24,000[1] species divided into about 20 families. Well-known or economically important members of this order include lavender, lilac, olive, jasmine, the ash tree, teak, snapdragon, sesame, psyllium, garden sage, and a number of table herbs such as mint, basil, and rosemary.
Description
Although exceptions occur, species in this order typically have the following characteristics:
- superior ovary composed of two fused carpels
- four petals fused into a tube
- bilaterally symmetrical, often bilabiate corollas
- four (or fewer) fertile stamens
Taxonomy
The Lamiales previously had a restricted circumscription (e.g., by Arthur Cronquist) that included the major families Lamiaceae (Labiatae), Verbenaceae, and Boraginaceae, plus a few smaller families. In the classification system of Dahlgren the Lamiales were in the superorder Lamiiflorae (also called Lamianae). Recent phylogenetic work has shown the Lamiales are polyphyletic with respect to order Scrophulariales and the two groups are now usually combined in a single order that also includes the former orders Hippuridales and Plantaginales. Lamiales has become the preferred name for this much larger combined group. The placement of the Boraginaceae is unclear, but phylogenetic work shows this family does not belong in Lamiales.
Also, the circumscription of family Scrophulariaceae, formerly a paraphyletic group defined primarily by plesiomorphic characters and from within which numerous other families of the Lamiales were derived, has been radically altered to create a number of smaller, better-defined, and putatively monophyletic families.
References
- 1 2 Angiosperm Phylogeny Group (2009). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III". Botanical Journal of the Linnean Society. 161 (2): 105–121. doi:10.1111/j.1095-8339.2009.00996.x.
- ↑ Angiosperm Phylogeny Group (2016). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG IV". Botanical Journal of the Linnean Society. 181 (1): 1–20. doi:10.1111/boj.12385.
External links
- Lamiales
- A parsimony analysis of the Asteridae sensu lato based on rbcL sequences
- Disintegration of the Scrophulariaceae (deals with relationships throughout Lamiales)
- L. Watson and M.J. Dallwitz (1992 onwards). The families of flowering plants: descriptions, illustrations, identification, information retrieval. http://delta-intkey.com
- http://www.biologie.uni-hamburg.de/b-online/vascular/acanth.htm 2002-09-06
- http://www.biologie.uni-hamburg.de/b-online/d52/52e.htm 2002-09-06
- http://www.biologie.uni-hamburg.de/b-online/d52/52efam.htm 2002-09-06
- http://www.science.siu.edu/parasitic-plants/Relation-Scroph.html
- http://www.rbgkew.org.uk/web.dbs/genlist.html 2002-09-06
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