Zopheridae

Zopheridae

Ironclad beetle, Zopherus nodulosus haldemani
Scientific classification
Kingdom:	Animalia
Subkingdom:	Eumetazoa
Phylum:	Arthropoda
Subphylum:	Hexapoda
Class:	Insecta
Order:	Coleoptera
Infraorder:	Cucujiformia
Superfamily:	Tenebrionoidea
Family:	Zopheridae Solier, 1834
Subfamilies
Colydiinae - cylindrical bark beetles Latometinae (sometimes in Zopheridae) Monommatinae (sometimes in Zopheridae) Pycnomerinae (sometimes in Zopheridae) Usechinae (sometimes in Zopheridae) Zopherinae - ironclad beetles

Zopheridae used as jewelry

The Zopheridae family of beetles has grown considerably in recent years as the members of two other families have been included within its circumscription; these former families are the Monommatidae and the Colydiidae, which are now both included in the Zopheridae as subfamilies or (in the former case) even as tribe of subfamily Zopherinae. Some authors accept up to six subfamilies here, while others merge all except the Colydiinae into the Zopherinae.

Over 100 genera are in the redefined family, and hundreds of species worldwide. No common name is used for the new family, though some of the constituent subfamilies have their own, including the ironclad beetles, and the cylindrical bark beetles.

Extant families	Amphizoidae (trout-stream beetles) Aspidytidae Carabidae (ground beetles) Dytiscidae (predaceous diving beetles) Gyrinidae (whirligig beetles) Haliplidae (crawling water beetles) Hygrobiidae Meruidae (Meru phyllisae) Noteridae (burrowing water beetles) Rhysodidae (wrinkled bark beetles) Trachypachidae (false ground beetles)

Suborder Myxophaga

Hydroscaphidae (skiff beetles) Lepiceridae Sphaeriusidae Torridincolidae

Suborder Polyphaga

Bostrichiformia

Bostrichoidea	Anobiidae (furniture beetles, death watch beetles, spider beetles) Bostrichidae (auger beetles) Dermestidae (skin beetles) Jacobsoniidae (Jacobson's beetles) Nosodendridae (wounded-tree beetles)

Derodontoidea	Derodontidae (tooth-necked fungus beetles)

Cucujiformia

Chrysomeloidea	Cerambycidae (longhorn beetles) Chrysomelidae (leaf beetles) Disteniidae Megalopodidae Orsodacnidae Oxypeltidae Vesperidae

Cleroidea	Acanthocnemidae (Acanthocnemus nigricans) Chaetosomatidae Cleridae (checkered beetles) Melyridae (soft-wing flower beetles) Phloiophilidae (Phloiophilus edwardsi) Phycosecidae Prionoceridae Trogossitidae (bark-gnawing beetles)

Cucujoidea	Alexiidae Biphyllidae (false skin beetles) Boganiidae Bothrideridae (dry bark beetles) Byturidae (fruitworm beetles) Cavognathidae Cerylonidae (minute bark beetles) Coccinellidae (lady beetles, or God's cows) Corylophidae (minute fungus beetles) Cryptophagidae (silken fungus beetles) Cucujidae (flat bark beetles) Cyclaxyridae Discolomatidae Endomychidae (handsome fungus beetles) Erotylidae (pleasing fungus beetles) Helotidae Hobartiidae Kateretidae (short-winged flower beetles) Laemophloeidae (lined flat bark beetles) Lamingtoniidae (Lamingtonium binnaberrense) Latridiidae (minute brown scavenger beetles) Monotomidae (root-eating beetles) Myraboliidae Nitidulidae (sap beetles) Passandridae (parasitic flat bark beetles) Phalacridae (shining flower beetles) Phloeostichidae Propalticidae Protocucujidae Silvanidae (silvanid flat bark beetles) Smicripidae (palmetto beetles) Sphindidae (dry-fungus beetles)

Curculionoidea (weevils)	Anthribidae (fungus weevils) Attelabidae (leaf-rolling weevils) Belidae (primitive weevils) Brentidae (straight snout weevils, New York weevil) Caridae Curculionidae (true weevils, bark beetles, ambrosia beetles) Nemonychidae (pine flower weevils)

Lymexyloidea	Lymexylidae (ship-timber beetles)

Tenebrionoidea	Aderidae (ant-like leaf beetles) Anthicidae (ant-like flower beetles) Archeocrypticidae (cryptic fungus beetles) Boridae (conifer bark beetles) Chalcodryidae Ciidae (minute tree-fungus beetles) Melandryidae (false darkling beetles) Meloidae (blister beetles) Mordellidae (tumbling flower beetles) Mycetophagidae (hairy fungus beetles) Mycteridae (palm and flower beetles) Oedemeridae (false blister beetle) Perimylopidae, or Promecheilidae Prostomidae (jugular-horned beetles) Pterogeniidae Pyrochroidae (fire-coloured beetles) Pythidae (dead log bark beetles) Ripiphoridae (wedge-shaped beetles) Salpingidae (narrow-waisted bark beetles) Scraptiidae (false flower beetles) Stenotrachelidae (false longhorn beetles) Synchroidae (synchroa bark beetles) Tenebrionidae (darkling beetles) Tetratomidae (polypore fungus beetles) Trachelostenidae Trictenotomidae Ulodidae Zopheridae (ironclad beetles, cylindrical bark beetles)

Elateriformia

Buprestoidea	Buprestidae (jewel beetles, or metallic wood-boring beetles) Schizopodidae

Byrrhoidea	Byrrhidae (pill beetles) Callirhipidae (cedar beetles) Chelonariidae (turtle beetles) Cneoglossidae Dryopidae (long-toed water beetles) Elmidae (riffle beetles) Eulichadidae (forest stream beetles) Heteroceridae (variegated mud-loving beetles) Limnichidae (minute mud beetles) Lutrochidae (travertine beetles) Psephenidae (water-penny beetles) Ptilodactylidae

Dascilloidea	Dascillidae (soft bodied plant beetles) Rhipiceridae (cicada beetle, cicada parasite beetles)

Elateroidea	Anischiidae Artematopodidae (soft-bodied plant beetles) Brachypsectridae (Texas beetles) Cantharidae (soldier beetles) Cerophytidae (rare click beetles) Drilidae Elateridae (click beetles) Eucnemidae (false click beetles) Lampyridae (fireflies) Lycidae (net-winged beetles) Omalisidae Omethidae (false fireflies) Phengodidae (glowworm beetles, long-lipped beetles) Plastoceridae (Plastocerus angulosus) Podabrocephalidae Rhagophthalmidae Rhinorhipidae (Rhinorhipus tamborinensis) Throscidae (false metallic wood-boring beetles)

Scirtoidea	Clambidae Decliniidae (Declinia relicta) Eucinetidae (plate-thigh beetles) Scirtidae

Scarabaeiformia

Scarabaeoidea

Staphyliniformia

Histeroidea	Histeridae (clown beetles) Sphaeritidae (false clown beetles) Synteliidae

Hydrophiloidea	Epimetopidae Georissidae (minute mud-loving beetles) Helophoridae Hydrochidae Hydrophilidae (water scavenger beetles) Spercheidae

Staphylinoidea	Agyrtidae (primitive carrion beetles) Hydraenidae Leiodidae (round fungus beetles) Ptiliidae (feather-winged beetles) Scydmaenidae (ant-like stone beetles) Silphidae (carrion beetles) Staphylinidae (rove beetles)

List of subgroups of the order Coleoptera

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Zopheridae

See also