2009 in arthropod paleontology

List of years in arthropod paleontology
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This list of fossil arthropods described in 2009 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods that have been described during the year 2009. The list only includes taxa at the level of genus or species.

Anomalocaridids

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Schinderhannes[1]	gen et sp nov	Valid	Kühl, Briggs, & Rust	Lower Devonian	Hunsrück Slate	Germany

Arachnids

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Palaeoperenethis[2]	gen et sp nov	Valid	Selden & Penney	Ypresian	Horsefly Lagerstätte, British Columbia	Canada

Insects

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Agulla mineralensis[3]	sp nov	Valid	Engel	Late Barstovian	Stewart Valley Group	USA	only described Neogene snakefly fossil
Allorapisma[4]	gen et sp nov	Valid	Makarkin & Archibald	Ypresian	Tom Thumb Tuff, Klondike Mountain Formation	USA		Allorapisma chuorum
Apis (Cascapis) nearctica[5]	sp nov	Valid	Engel, Hinojosa-Diaz, & Rasnitsyn	Middle Miocene		USA	The first fossil Apis from North America.
Aspidopleura[6]	gen et sp nov	Valid	Gibson	Eocene	Baltic amber			Aspidopleura baltica
Brevivulva[6]	gen et sp nov	Valid	Gibson	Eocene	Baltic amber			Brevivulva electroma
Cimbrophlebia brooksi[7]	sp nov	Valid	Archibald	Ypresian	Tom Thumb Tuff, Klondike Mountain Formation	USA	A Cimbrophlebiid scorpionfly
Cimbrophlebia flabelliformis[7]	sp nov	Valid	Archibald	Ypresian	Kamloops Group, McAbee, British Columbia	Canada
Cimbrophlebia leahyi[7]	sp nov	Valid	Archibald	Ypresian	Kamloops Group, McAbee, British Columbia	Canada
Cimbrophlebia westae[7]	sp nov	Valid	Archibald	Ypresian	Tom Thumb Tuff, Klondike Mountain Formation	USA
Denaeaspis[8]	gen et sp nov	valid	Chaboo & Engel	Lutetian	Parachute Member, Green River Formation	USA	One of the oldest tortoise beetles
Eosacantha[8]	gen et sp nov	valid	Chaboo & Engel	Lutetian	Parachute Member, Green River Formation	USA	One of the oldest tortoise beetles
Gesomyrmex germanicus[9]	sp nov	Valid	Dlussky, Wappler, & Wedmann	Lutetian	Eckfeld Maar	Germany	A Formicinae ant	Gesomyrmex germanicus
Gesomyrmex pulcher[9]	sp nov	Valid	Dlussky, Wappler, & Wedmann	Lutetian	Messel Formation	Germany	A Formicinae ant	Gesomyrmex pulcher
Leptofoenus pittfieldae[10]	sp nov	Valid	Engel	Burdigalian	Dominican amber	Dominican Republic		Leptofoenus pittfieldae
Metapelma archetypon[6]	sp nov	Valid	Gibson	Eocene	Baltic amber			Metapelma archetypon
Neanaperiallus[6]	gen et sp nov	Valid	Gibson	Eocene	Baltic amber			Neanaperiallus masneri
Nesagapostemon[11]	gen et sp nov	Valid	Engel	Burdigalian	Dominican amber	Dominican Republic		Nesagapostemon moronei
Nymphes georgei[12]	sp nov	Valid	Archibald, Makarkin, & Ansorge	Ypresian	Tom Thumb Tuff, Klondike Mountain Formation	USA		Nymphes georgei
Oligochlora semirugosa[11]	sp nov	Valid	Engel	Burdigalian	Dominican amber	Dominican Republic		Oligochlora semirugosa
Principiala rudgwickensis[13]	sp nov	valid	Jepson, Makarkin, & Jarzembowski	Barremian	Upper Weald Clay	England	An Ithonidae lacewing, a second species of Principiala
Proceratium eocenicum[14]	Sp nov	Valid	Dlussky	Late Eocene	Baltic amber	Russia	A ponerine ant	Proceratium eocenicum
Pronymphes hoffeinsorum[12]	Sp nov	Valid	Archibald, Makarkin, Ansorge	Priabonian	Yantarny [=Palmnicken], Kaliningradskaya Oblast’	Russia
Termitaradus mitnicki[15]	sp nov	Valid	Engel	Burdigalian	Dominican amber	Dominican Republic	A termite bug	Termitaradus mitnicki

References

1. Kühl, G; Briggs, DE; Rust, J (February 2009). "A Great-Appendage Arthropod with a Radial Mouth from the Lower Devonian Hunsrück Slate, Germany". Science. 323 (5915): 771–3. Bibcode:2009Sci...323..771K. doi:10.1126/science.1166586. ISSN 0036-8075. PMID 19197061.
2. Selden, P.A.; Penney, D. (2009). "A fossil spider (Araneae: Pisauridae) of Eocene age from Horsefly, British Columbia, Canada" (PDF). Contributions to Natural History. 12: 1269–1282.
3. Engel, M. S. (2009). "A Miocene snakefly from Stewart Valley, Nevada (Raphidioptera: Raphidiidae)". Transactions of the Kansas Academy of Science. 112 (4): 211–214. doi:10.1660/062.112.0409.
4. Makarkin, V.N.; Archibald, S.B. (2009). "A new genus and first Cenozoic fossil record of moth lacewings (Neuroptera: Ithonidae) from the Early Eocene of North America" (PDF). Zootaxa. 2063: 55–63. Retrieved 24 March 2010.
5. Michael S. Engel, I. A. Hinojosa-Diaz & A. P. Rasnitsyn (2009). "A honey bee from the Miocene of Nevada and the biogeography of Apis (Hymenoptera: Apidae: Apini)" (PDF). Proceedings of the California Academy of Sciences. 60 (3): 23–38. Retrieved 4 September 2010.
6. Gibson, G.A.P. (2009). "Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa". ZooKeys. 20: 175–214. doi:10.3897/zookeys.20.161. Retrieved 24 March 2010.
7. Archibald, S.B. (2009). "New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA" (PDF). Zootaxa. 2249: 51–62. Retrieved 24 March 2010.
8. Chaboo, C. S.; Engel, M. S. (2009). "Eocene tortoise beetles from the Green River Formation in Colorado, U.S.A. (Coleoptera: Chrysomelidae: Cassidinae)". Systematic Entomology. 34 (2): 202–209. doi:10.1111/j.1365-3113.2008.00456.x.
9. Dlussky, GM; Wappler, T; Wedmann, S (2009). "Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities". Zootaxa. 2031: 1–20.
10. Engel, M.S. (2005). "The first fossil leptofoenine wasp (Hymenoptera, Pteromalidae): A new species of Leptofoenusin Miocene amber from the Dominican Republic". ZooKeys. 13: 57–66. doi:10.3897/zookeys.13.159.
11. Engel, M.S. (2009). "Two new halictine bees in Miocene Amber from the Dominican Republic (Hymenoptera, Halictidae)". ZooKeys. 29: 1–12. doi:10.3897/zookeys.29.257.
12. Archibald, S.B.; Makarkin,V.N.; Ansorge, J. (2009). "New fossil species of Nymphidae (Neuroptera) from the Eocene of North America and Europe" (PDF). Zootaxa. 2157: 59–68.
13. Jepson, JE; Makarkin, VN; Jarzembowski, E (2009). "New lacewings (Insecta: Neuroptera) from the Lower Cretaceous Wealden supergroup of Southern England". Cretaceous Research. 30: 1325–1338. doi:10.1016/j.cretres.2009.07.012.
14. Dlussky, G. M. (2009). "The ant subfamilies Ponerinae, Cerapachyinae, and Pseudomyrmecinae (Hymenoptera, Formicidae) in the Late Eocene Ambers of Europe". Paleontological Journal. 43: 1043–1086.
15. Engel, M.S. (2009). "A new termite bug in Miocene amber from the Dominican Republic (Hemiptera, Termitaphididae)". ZooKeys. 25: 61–68. doi:10.3897/zookeys.25.267.