List of butterflies of Sudan and South Sudan

Location of Sudan

This is a list of butterflies of Sudan and South Sudan. About 341 species are known from the Sudans,^[1] 6 of which are endemic.^[2]

Papilionidae

Papilioninae

Papilionini

Papilio antimachus Drury, 1782
Papilio zalmoxis Hewitson, 1864
Papilio nireus lyaeus Doubleday, 1845
Papilio nireus pseudonireus Felder & Felder, 1865
Papilio plagiatus Aurivillius, 1898
Papilio phorcas sudanicola Storace, 1965
Papilio rex franciscae Carpenter, 1928
Papilio mechowi Dewitz, 1881
Papilio mechowianus Dewitz, 1885
Papilio echerioides joiceyi Gabriel, 1945
Papilio jacksoni imatonga Clifton & Collins, 1997
Papilio nobilis didingensis Carpenter, 1928
Papilio hesperus sudana Gabriel, 1945
Papilio lormieri Distant, 1874
Papilio mackinnoni Sharpe, 1891

Leptocercini

Graphium antheus (Cramer, 1779)
Graphium policenes policenes (Cramer, 1775)
Graphium policenes telloi Hecq, 1999
Graphium colonna (Ward, 1873)
Graphium angolanus baronis (Ungemach, 1932)
Graphium ridleyanus (White, 1843)
Graphium leonidas (Fabricius, 1793)
Graphium philonoe whalleyi (Talbot, 1929)
Graphium almansor escherichi (Gaede, 1915)

Pieridae

Lycaenidae

Miletinae

Miletini

Megalopalpus zymna (Westwood, 1851)
Lachnocnema abyssinica Libert, 1996
Lachnocnema emperamus (Snellen, 1872)
Lachnocnema divergens Gaede, 1915
Lachnocnema reutlingeri perspicua Libert, 1996

Lipteninae

Liptenini

Alaena subrubra Bethune-Baker, 1915
Pentila pauli multiplagata Bethune-Baker, 1908
Pentila umangiana connectens Hulstaert, 1924
Telipna sulpitia Hulstaert, 1924
Telipna albofasciata laplumei Devos, 1917
Ornipholidotos overlaeti intermedia Libert, 2005
Ornipholidotos latimargo (Hawker-Smith, 1933)
Mimacraea krausei karschioides Carpenter & Jackson, 1950
Liptena ferrymani (Grose-Smith & Kirby, 1891)
Liptena xanthostola xantha (Grose-Smith, 1901)
Tetrarhanis ilala (Riley, 1929)
Larinopoda lircaea (Hewitson, 1866)

Epitolini

Stempfferia cercenoides (Holland, 1890)
Cephetola dolorosa (Roche, 1954)
Phytala elais ugandae Jackson, 1964

Theclinae

Myrina silenus (Fabricius, 1775)
Myrina subornata nuba Talbot, 1935
Dapidodigma demeter sudsudana d'Abrera, 1980

Aphnaeini

Pseudaletis agrippina Druce, 1888
Chloroselas tamaniba (Walker, 1870)
Chloroselas taposana Riley, 1932
Cigaritis acamas bellatrix (Butler, 1886)
Cigaritis nilus (Hewitson, 1865)
Zeritis neriene Boisduval, 1836
Zeritis pulcherrima Aurivillius, 1923
Axiocerses harpax kadugli Talbot, 1935
Axiocerses amanga (Westwood, 1881)
Aphnaeus coronae Talbot, 1935

Theclini

Hypolycaena liara Druce, 1890
Hypolycaena obscura Stempffer, 1947
Leptomyrina sudanica Stempffer, 1964
Iolaus alienus ugandae Stempffer, 1953
Iolaus scintillans Aurivillius, 1905
Iolaus sudanicus Aurivillius, 1905
Iolaus pallene (Wallengren, 1857)
Iolaus menas Druce, 1890
Iolaus ismenias (Klug, 1834)
Stugeta marmoreus (Butler, 1866)
Paradeudorix ituri ugandae (Talbot, 1935)
Hypomyrina mimetica Libert, 2004
Deudorix dinochares Grose-Smith, 1887
Deudorix dinomenes diomedes Jackson, 1966
Deudorix livia (Klug, 1834)
Deudorix lorisona baronica Ungemach, 1932
Capys bamptoni Henning & Henning, 1988

Polyommatinae

Lycaenesthini

Anthene butleri (Oberthür, 1880)
Anthene crawshayi minuta (Bethune-Baker, 1916)
Anthene hodsoni (Talbot, 1935)
Anthene lunulata (Trimen, 1894)
Anthene wilsoni (Talbot, 1935)
Anthene lusones (Hewitson, 1874)
Anthene nigeriae (Aurivillius, 1905)

Polyommatini

Cupidopsis jobates mauritanica Riley, 1932
Uranothauma delatorum Heron, 1909
Uranothauma heritsia intermedia (Tite, 1958)
Leptotes marginalis (Stempffer, 1944)
Tuxentius margaritaceus (Sharpe, 1892)
Tarucus balkanicus (Freyer, 1843)
Tarucus legrasi Stempffer, 1948
Tarucus rosacea (Austaut, 1885)
Tarucus theophrastus (Fabricius, 1793)
Tarucus ungemachi Stempffer, 1942
Zizeeria karsandra (Moore, 1865)
Actizera stellata (Trimen, 1883)
Euchrysops alberta (Butler, 1901)
Euchrysops albistriata (Capronnier, 1889)
Euchrysops crawshayi (Butler, 1899)
Euchrysops cyclopteris (Butler, 1876)
Euchrysops nilotica (Aurivillius, 1904)
Chilades eleusis (Demaison, 1888)
Lepidochrysops albilinea Tite, 1959
Lepidochrysops negus wau (Wichgraf, 1921)
Lepidochrysops nigrita Tite, 1959
Lepidochrysops polydialecta (Bethune-Baker, [1923])
Lepidochrysops victoriae occidentalis Libert & Collins, 2001

Nymphalidae

Danainae

Danaini

Danaus chrysippus chrysippus (Linnaeus, 1758)
Danaus chrysippus alcippus (Cramer, 1777)
Danaus dorippus (Klug, 1845)
Tirumala formosa neumanni (Rothschild, 1902)
Tirumala petiverana (Doubleday, 1847)
Amauris niavius aethiops Rothschild & Jordan, 1903
Amauris tartarea Mabille, 1876
Amauris albimaculata sudanica Talbot, 1940
Amauris echeria mongallensis Carpenter, 1928
Amauris echeria steckeri Kheil, 1890
Amauris hecate (Butler, 1866)

Satyrinae

Melanitini

Gnophodes betsimena parmeno Doubleday, 1849
Melanitis libya Distant, 1882

Satyrini

Bicyclus angulosa (Butler, 1868)
Bicyclus campus (Karsch, 1893)
Bicyclus istaris (Plötz, 1880)
Bicyclus kenia (Rogenhofer, 1891)
Bicyclus milyas (Hewitson, 1864)
Bicyclus pavonis (Butler, 1876)
Bicyclus martius sanaos (Hewitson, 1866)
Bicyclus saussurei angustus Condamin, 1970
Heteropsis perspicua (Trimen, 1873)
Ypthima albida Butler, 1888
Ypthima antennata van Son, 1955
Ypthima condamini Kielland, 1982
Ypthima doleta Kirby, 1880
Ypthima pupillaris Butler, 1888
Ypthima simplicia Butler, 1876
Ypthimomorpha itonia (Hewitson, 1865)

Charaxinae

Charaxini

Charaxes varanes vologeses (Mabille, 1876)
Charaxes fulvescens monitor Rothschild, 1900
Charaxes boueti rectans Rothschild & Jordan, 1903
Charaxes cynthia kinduana Le Cerf, 1923
Charaxes lactetinctus ungemachi Le Cerf, 1927
Charaxes jasius Poulton, 1926
Charaxes epijasius Reiche, 1850
Charaxes hansali baringana Rothschild, 1905
Charaxes brutus angustus Rothschild, 1900
Charaxes julius somalicus Rothschild, 1900
Charaxes ansorgei kinyeti Plantrou, 1989
Charaxes pollux (Cramer, 1775)
Charaxes druceanus vivianae Plantrou, 1982
Charaxes eudoxus imatongensis Plantrou, 1982
Charaxes eudoxus mechowi Rothschild, 1900
Charaxes numenes aequatorialis van Someren, 1972
Charaxes tiridates tiridatinus Röber, 1936
Charaxes bipunctatus ugandensis van Someren, 1972
Charaxes smaragdalis gobyae Plantrou, 1989
Charaxes etesipe (Godart, 1824)
Charaxes achaemenes monticola van Someren, 1970
Charaxes jahlusa ganalensis Carpenter, 1937
Charaxes eupale latimargo Joicey & Talbot, 1921
Charaxes dilutus ngonga van Someren, 1974
Charaxes baumanni didingensis van Someren, 1971
Charaxes etheocles carpenteri van Someren & Jackson, 1957
Charaxes cedreatis Hewitson, 1874
Charaxes viola picta van Someren & Jackson, 1952
Charaxes paphianus subpallida Joicey & Talbot, 1925
Charaxes kahldeni Homeyer & Dewitz, 1882
Charaxes zoolina (Westwood, [1850])
Charaxes lycurgus bernardiana Plantrou, 1978
Charaxes zelica depuncta Joicey & Talbot, 1921
Charaxes amandae Rydon, 1989

Euxanthini

Charaxes crossleyi ansorgei (Rothschild, 1903)

Apaturinae

Apaturopsis cleochares (Hewitson, 1873)

Nymphalinae

Nymphalini

Antanartia dimorphica Howarth, 1966
Junonia chorimene (Guérin-Méneville, 1844)
Junonia terea tereoides (Butler, 1901)
Junonia westermanni Westwood, 1870
Protogoniomorpha parhassus (Drury, 1782)
Protogoniomorpha temora (Felder & Felder, 1867)
Precis archesia ugandensis (McLeod, 1980)
Precis coelestina Dewitz, 1879
Precis octavia (Cramer, 1777)
Precis tugela aurorina (Butler, 1894)
Hypolimnas anthedon (Doubleday, 1845)
Hypolimnas misippus (Linnaeus, 1764)
Hypolimnas salmacis platydema Rothschild & Jordan, 1903
Mallika jacksoni (Sharpe, 1896)

Biblidinae

Biblidini

Byblia anvatara acheloia (Wallengren, 1857)
Mesoxantha ethosea reducta Rothschild, 1918
Neptidopsis ophione nucleata Grünberg, 1911

Limenitinae

Limenitidini

Pseudoneptis bugandensis Stoneham, 1935
Pseudacraea acholica Riley, 1932
Pseudacraea boisduvalii (Doubleday, 1845)
Pseudacraea eurytus (Linnaeus, 1758)
Pseudacraea lucretia protracta (Butler, 1874)

Neptidini

Neptis metella (Doubleday, 1848)
Neptis nemetes nemetes Hewitson, 1868
Neptis nemetes margueriteae Fox, 1968
Neptis occidentalis Rothschild, 1918

Adoliadini

Catuna crithea (Drury, 1773)
Pseudargynnis hegemone (Godart, 1819)
Euphaedra medon fraudata van Someren, 1935
Euphaedra neumanni Rothschild, 1902
Euphaedra harpalyce sudanensis Talbot, 1929

Heliconiinae

Acraeini

Acraea chilo Godman, 1880
Acraea insignis Distant, 1880
Acraea neobule Doubleday, 1847
Acraea pseudolycia astrigera Butler, 1899
Acraea quirina (Fabricius, 1781)
Acraea zetes (Linnaeus, 1758)
Acraea abdera Hewitson, 1852
Acraea cepheus (Linnaeus, 1758)
Acraea egina (Cramer, 1775)
Acraea guluensis Le Doux, 1932
Acraea caecilia (Fabricius, 1781)
Acraea caldarena Hewitson, 1877
Acraea doubledayi Guérin-Méneville, 1849
Acraea marnois Rogenhofer, 1890
Acraea pseudegina Westwood, 1852
Acraea sykesi Sharpe, 1902
Acraea aganice orientalis (Ungemach, 1932)
Acraea epaea paragea (Grose-Smith, 1900)
Acraea epiprotea (Butler, 1874)
Acraea macarista (Sharpe, 1906)
Acraea poggei nelsoni Grose-Smith & Kirby, 1892
Acraea pseuderyta Godman & Salvin, 1890
Acraea acerata Hewitson, 1874
Acraea bonasia (Fabricius, 1775)
Acraea encedana Pierre, 1976
Acraea serena (Fabricius, 1775)
Acraea esebria Hewitson, 1861
Acraea jodutta (Fabricius, 1793)
Acraea johnstoni Godman, 1885
Acraea lycoa Godart, 1819
Acraea pharsalus pharsalus Ward, 1871
Acraea pharsalus rhodina Rothschild, 1902
Acraea oreas Sharpe, 1891

Argynnini

Issoria hanningtoni imatonga (Riley, 1932)

Vagrantini

Lachnoptera anticlia (Hübner, 1819)
Phalanta eurytis eurytis (Doubleday, 1847)
Phalanta eurytis microps (Rothschild & Jordan, 1903)
Phalanta phalantha aethiopica (Rothschild & Jordan, 1903)

Hesperiidae

Coeliadinae

Coeliades forestan (Stoll, [1782])
Coeliades keithloa menelik (Ungemach, 1932)
Coeliades pisistratus (Fabricius, 1793)

Pyrginae

Celaenorrhinini

Celaenorrhinus proxima (Mabille, 1877)
Eretis lugens (Rogenhofer, 1891)
Eretis melania Mabille, 1891
Eretis mixta Evans, 1937
Sarangesa aza Evans, 1951
Sarangesa brigida sanaga Miller, 1964
Sarangesa laelius (Mabille, 1877)
Sarangesa lucidella (Mabille, 1891)
Sarangesa phidyle (Walker, 1870)

Tagiadini

Tagiades flesus (Fabricius, 1781)
Eagris decastigma purpura Evans, 1937
Eagris denuba obliterata Carpenter, 1928
Eagris lucetia (Hewitson, 1875)
Eagris tigris liberti Collins & Larsen, 2005
Caprona adelica Karsch, 1892
Netrobalane canopus (Trimen, 1864)
Abantis bismarcki Karsch, 1892

Carcharodini

Spialia doris (Walker, 1870)
Spialia zebra bifida (Higgins, 1924)

Hesperiinae

Aeromachini

Prosopalpus styla Evans, 1937
Kedestes protensa Butler, 1901
Kedestes rogersi Druce, 1907
Gorgyra aretina (Hewitson, 1878)
Pardaleodes incerta (Snellen, 1872)
Pardaleodes sator pusiella Mabille, 1877
Pardaleodes tibullus (Fabricius, 1793)
Osmodes thora (Plötz, 1884)
Acleros mackenii olaus (Plötz, 1884)
Andronymus caesar philander (Hopffer, 1855)
Chondrolepis niveicornis (Plötz, 1883)
Zophopetes dysmephila (Trimen, 1868)
Gretna cylinda (Hewitson, 1876)
Caenides dacena (Hewitson, 1876)
Fresna nyassae (Hewitson, 1878)

Baorini

Borbo fallax (Gaede, 1916)
Borbo perobscura (Druce, 1912)
Gegenes nostrodamus (Fabricius, 1793)

Heteropterinae

Metisella quadrisignatus (Butler, 1894)
Metisella tsadicus (Aurivillius, 1905)
Lepella lepeletier (Latreille, 1824)

References

List of butterflies of Africa

Sovereign states	Algeria Angola Benin Botswana Burkina Faso Burundi Cameroon Cape Verde Central African Republic Chad Comoros Democratic Republic of the Congo Republic of the Congo Djibouti Egypt Equatorial Guinea Eritrea Ethiopia Gabon The Gambia Ghana Guinea Guinea-Bissau Ivory Coast (Côte d'Ivoire) Kenya Lesotho Liberia Libya Madagascar Malawi Mali Mauritania Mauritius Morocco Mozambique Namibia Niger Nigeria Rwanda São Tomé and Príncipe Senegal Seychelles Sierra Leone Somalia South Africa South Sudan Sudan Swaziland Tanzania Togo Tunisia Uganda Zambia Zimbabwe

States with limited recognition	Sahrawi Arab Democratic Republic Somaliland

Dependencies and other territories	Canary Islands / Ceuta / Melilla / Plazas de soberanía (Spain) Madeira (Portugal) Mayotte / Réunion (France) Saint Helena / Ascension Island / Tristan da Cunha (United Kingdom)

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