List of coleopterans of Sri Lanka

Sri Lanka is a tropical island situated close to the southern tip of India. The invertebrate fauna is as large as it is common to other regions of the world. There are about 2 million species of arthropods found in the world, and still it is counting. So many new species are discover up to this time also. So it is very complicated and difficult to summarize the exact number of species found within a certain region.

This a list of the coleopterans found from Sri Lanka.

Beetle

Phylum: Arthropoda
Class: Insecta
Order: Coleoptera

Beetles are a group of insects that form the order Coleoptera. They have two pairs of wings, the front pair, the "elytra", being hardened and thickened into a shell-like protection for the rear pair and the beetle's abdomen. The order contains more species than any other order, constituting almost 25% of all known animal life-forms. About 40% of all described insect species are beetles (about 400,000 species), and new species are discovered frequently. The largest taxonomic family, the Curculionidae (the weevils or snout beetles), also belongs to this order.

The diversity of beetles is very wide-ranging. They are found in almost all types of habitats, but are not known to occur in the sea or in the polar regions. In Sri Lanka, particular beetle families are extensively studied,[1][2] but some others not at all.[3] Most of the studies beetles are due to the fact that they are pests for many cultivations.[4] Numerous works had been done in the 1980s and 1990s by foreign scientists such as Arrow (1910-1931), Franz (1982), Hammond (1972), Hansen (1999), Chatterjee (1924) and Bonadona (1986). The knowledge by local investigators opened in the early 2000s and in 2003, Prof. Dangalle studies heavily on tiger beetles. Then in 2013, Dr. Qader et al. studies beetles in Western province to find their ecological niches and distribution.[5] Weevils were studied by Prof. Gunawardena since 1993 and predatory Coccinellid beetles were studies by Dr. S Mayadunnage.[6][7]

According to Channa Bambaradeniya et al., 3,033 species of coleopterans belongs to 115 families[8] are recorded from Sri Lanka, which comprises the largest animal group in Sri Lanka.[9]

Family: Anthicidae - Ant-like beetles

  • Amblyderus bigibber
  • Amblyderus brincki
  • Anthelephila consul
  • Anthelephila insulana
  • Anthicus semirubidus
  • Formicomus anderssoni
  • Formicomus braminus
  • Formicomus opiatus
  • Leptaleus brincki
  • Mecynotarsus cederholmi
  • Mecynotarsus flinti
  • Notoxus brinckianus
  • Notoxus ravana
  • Omonadus formicarius
  • Pseudonotoxus cederholmi
  • Pseudonotoxus minutus
  • Sapintus barbei
  • Sapintus brincki
  • Sapintus capitatus
  • Sapintus flinti
  • Tomoderus mussardi

Family: Anthribidae - Fungus weevils

  • Acorynus ceylonicus
  • Acorynus cinereomaculatus
  • Acorynus cingalus
  • Acorynus dohrni
  • Acorynus labidus
  • Acorynus passerinus
  • Apolecta nietneri
  • Araecerus fragilis
  • Araecerus intangens
  • Araecerus irresolutus
  • Araecerus pardalis
  • Araecerus pumilus
  • Atinellia senex
  • Basitropis nitidicutis
  • Caccorhinus brunnipennis
  • Caccorhinus disconotatus
  • Contexta murina
  • Dendrotrogus colligens
  • Dendrotrogus hypocrita
  • Directarius bifoveatus
  • Disphaerona picta
  • Disphaerona punctata
  • Disphaerona verrucosus
  • Exechesops horni
  • Exechesops molitor
  • Habrissus asellus
  • Habrissus tibialis
  • Hybosternus mesosternalis
  • Hypseus dilectus
  • Litocerus annulipes
  • Litocerus crucicollis
  • Litocerus semiustus
  • Mauia subnotata
  • Mecotropis bipunctatus
  • Mecotropis gardneri
  • Melanopsacus ceylonicus
  • Misthosima separ
  • Phaulimia caena
  • Phaulimia disticha
  • Phaulimia schaumi
  • Phloeobius alternans
  • Phloeobius ceylonicus
  • Rhaphitropis tamilis
  • Straboscopus revocans
  • Straboscopus riehli
  • Sympaector angulifer
  • Tropideres verrucosus
  • Tropidobasis gemella
  • Ulorhinus distichus
  • Uncifer diffinis
  • Urodon maculatus
  • Urodon nigripes
  • Urodon tantillus
  • Xenocerus khasianus
  • Xenocerus mesosternalis
  • Xenocerus rectilineatus
  • Xylinada tamilanus
  • Xylinada indigus
  • Zygaenodes horni
  • Zygaenodes molitor

Family: Aphodiidae - Aphodiine dung beetles

  • Aganocrossus amoenus
  • Aganocrossus pallidicornis
  • Alocoderus elongatulus
  • Ammoecioides spectabilis - ssp. tricarinulatus
  • Aphodius humilis
  • Aphodius lewisi
  • Aphodius urostigma
  • Ataenius ceylonensis
  • Calamosternus granarius
  • Chaetopisthes singalensis
  • Labarrus hoabinhensis
  • Leiopsammodius indicus
  • Liothorax kraatzi
  • Mesontoplatys rufolaterus
  • Neocalaphodius moestus
  • Neotrichiorhyssemus expansicollis
  • Neotrichiorhyssemus hegeri
  • Paradidactylia avunculus
  • Paradidactylia biseriatus
  • Paradidactylia carinulatus
  • Paradidactylia haafi
  • Paradidactylia ovatulus
  • Paradidactylia wichei
  • Phalacronothus ceylonensis
  • Pharaphodius crenatus
  • Pharaphodius robustus
  • Pharaphodius lewisi
  • Pharaphodius ratnapurensis
  • Pleurophorus cracens
  • Psammodius tesari
  • Rhyssemus granulosus
  • Rhyssemus indicus
  • Rhyssemus inscitus
  • Rhyssemus loebli
  • Saprosites dynastoides
  • Saprosites loebli
  • Saprosites sulciceps
  • Sybax impressicollis
  • Trichaphodius moorei
  • Trichiorhyssemus fruhstorferi

Family: Attelabidae - Leaf-rolling weevils

  • Apoderus dohrni
  • Apoderus hystrix
  • Apoderus nietneri
  • Apoderus pulchellus
  • Apoderus pullus
  • Apoderus scitulus
  • Apoderus tranquebaricus
  • Apoderus verrucosus
  • Attelabus hystrix
  • Attelabus octospilotus
  • Attelabus tranquebaricus
  • Centrocorynus dohrni
  • Centrocorynus pulchellus
  • Euops metallica
  • Euops nietneri
  • Euops suffundens
  • Henicolabus octospilotus
  • Hoplapoderus echinatus
  • Hoplapoderus hystrix
  • Hoplapoderus nepalensis
  • Paracycnotrachelus nietneri

Family: Bostrichidae - Auger beetles

Family: Bothrideridae - Cocoon-forming beetles

Family: Brachyceridae - Tertiary-brachycerid weevils

  • Amblyrrhinus posticus
  • Desmidophorus caelatus
  • Desmidophorus communicans
  • Desmidophorus discriminans
  • Desmidophorus fasciculicollis
  • Desmidophorus inexpertus
  • Desmidophorus obliquefasciatus
  • Desmidophorus strenuus
  • Phytonomus ochraceus

Family: Brentidae - Straight-snouted weevils

Family: Bupestidae - Jewel beetles

Family: Cantharidae - Soldier beetle

Family: Carabidae - Ground beetles

  • Abacetus antiquus
  • Abacetus atratus
  • Abacetus bipunctatus
  • Abacetus cordicollis
  • Abacetus dejeani
  • Abacetus flavipes
  • Abacetus guttula
  • Abacetus pallipes
  • Abacetus pomptus
  • Abacetus rufopiceus
  • Abacetus semimetallicus
  • Abacetus testaceipes
  • Abacidus atratus
  • Acupalpus derogatus
  • Acupalpus sinuellus
  • Agonum (Agonum) chinense
  • Agonum illocatum
  • Agonum japonicum
  • Amblystomus bivittatus
  • Amblystomus femoralis
  • Amblystomus fuscescens
  • Amblystomus guttatus
  • Amblystomus guttula
  • Amblystomus indicus
  • Amblystomus mandibularis
  • Amblystomus punctatus
  • Amblystomus quadriguttatus
  • Amblystomus stenolophoides
  • Amblystomus vulneratus
  • Anaulacus adelioides
  • Anaulacus fasciatus - ssp. fasciatus
  • Anaulacus opaculus
  • Anaulacus pleuronectes
  • Anchista brunnea
  • Anomotachys acaroides
  • Anomotarus decoratus
  • Anomotarus stigmula
  • Anthracus anamensis
  • Anthracus horni
  • Apotomus atripennis
  • Apristus cupreus
  • Apristus subtransparens
  • Arame macra
  • Archicolliuris bimaculata - ssp. bimaculata
  • Archicolliuris immaculata
  • Ardistomopsis myrmex
  • Ardistomopsis ovicollis
  • Argiloborus amblygonus
  • Argiloborus ceylanicus
  • Argiloborus curtus
  • Argiloborus monticola
  • Argiloborus stricticollis
  • Arrowina pygmaea
  • Arrowina taprobanae
  • Batoscelis oblonga
  • Bembidion foveolatum
  • Bembidion opulentum - ssp. opulentum
  • Brachinus immaculicornis - ssp. immaculicornis
  • Brachinus limbellus
  • Brachinus pictus
  • Brachinus sexmaculatus
  • Brachinus tetracolon
  • Brachyodes subolivaceus
  • Brachyodes virens
  • Bradybaenus festivus
  • Caelostomus picipes
  • Caelostomus sculptipennis
  • Calleida splendidula
  • Callistomimus littoralis - ssp. ceylonicus
  • Callistomimus pernix
  • Callytron limosum
  • Catascopus cingalensis
  • Catascopus whithillii
  • Cerapterus latipes
  • Chlaeminus biguttatus
  • Chlaenius bengalensis
  • Chlaenius bioculatus
  • Chlaenius circumdatus - ssp. circumdatus
  • Chlaenius cyanostolus
  • Chlaenius fastigatus
  • Chlaenius fletcheri
  • Chlaenius hamifer
  • Chlaenius henryi
  • Chlaenius inops
  • Chlaenius kerkvoordeae
  • Chlaenius laetiusculus
  • Chlaenius laetus
  • Chlaenius lafertei
  • Chlaenius leucops
  • Chlaenius macropus
  • Chlaenius malachinus
  • Chlaenius nepalensis
  • Chlaenius nigricans
  • Chlaenius oodioides
  • Chlaenius orbicollis
  • Chlaenius parallelus
  • Chlaenius pretiosus
  • Chlaenius pulcher
  • Chlaenius punctatostriatus
  • Chlaenius ruficauda
  • Chlaenius rufifemoratus
  • Chlaenius rugulosus
  • Chlaenius scapularis
  • Chlaenius tetragonoderus - ssp. tetragonoderus
  • Chlaenius trachys
  • Chlaenius velocipes
  • Chlaenius xanthospilus
  • Cicindela aurulenta - ssp. juxtata
  • Cicindela bicolor - ssp. haemorrhoidalis, xanthospilota
  • Cicindela calligramma
  • Cicindela cardoni
  • Cicindela ceylonensis
  • Cicindela discrepans
  • Cicindela diversa
  • Cicindela fischeri - ssp. fischeri
  • Cicindela flavomaculata - ssp. flavomaculata
  • Cicindela lacrymans
  • Cicindela ocellata
  • Cicindela separata
  • Cicindela severa
  • Cicindela sylvicola
  • Clivina castanea
  • Clivina elongatula
  • Clivina forcipata
  • Clivina mustela
  • Clivina obenbergeri
  • Clivina striata
  • Clivina tranquebarica
  • Clivina westwoodi
  • Coleolissus iris
  • Collyris dohrnii - ssp. dohrnii
  • Colpodes bipars
  • Colpodes fletcheri
  • Colpodes iteratus
  • Colpodes repletus
  • Colpodes retusus
  • Colpodes sebosus
  • Colpodes xenos
  • Coptodera chaudoiri - ssp. chaudoiri
  • Coptodera eluta
  • Coptodera interrupta
  • Coptolobus anodon
  • Coptolobus ater
  • Coptolobus glabriculus
  • Coptolobus latus
  • Coptolobus lucens
  • Coptolobus omodon
  • Coryza maculata
  • Craspedophorus angulatus
  • Craspedophorus bifasciatus
  • Craspedophorus elegans
  • Craspedophorus halyi
  • Craspedophorus microspilotus
  • Creagris labrosa
  • Crepidogaster ceylanica
  • Crepidogaster horni
  • Cyclosomus flexuosus
  • Cylindera dormeri
  • Cylindera ganglbaueri
  • Cylindera henryi
  • Cylindera labioaenea - ssp. labioaenea
  • Cylindera lacunosa
  • Cylindera nietneri
  • Cylindera paradoxa
  • Cylindera singalensis
  • Cylindera waterhousei
  • Cylindera willeyi
  • Cymindoidea munda
  • Derocrania agnes
  • Derocrania concinna
  • Derocrania flavicornis
  • Derocrania fusiformis
  • Derocrania glabiceps
  • GleneDerocrania halyi
  • Derocrania intricatorugulosa
  • Derocrania jaechi
  • Derocrania nematodes
  • Derocrania nietneri
  • Derocrania shaumi
  • Derocrania scitiscabra
  • Desera geniculata
  • Dicaelindus impunctatus
  • Diceromerus orientalis
  • Dicranoncus cinctipennis
  • Dicranoncus queenslandicus
  • Dicranoncus ravus
  • Dioryche chinnada
  • Dioryche colombensis
  • Dioryche sericea
  • Dioryche torta - ssp. torta
  • Diplocheila daldorfi
  • Diplocheila laevis
  • Diplocheila polita
  • Dischissus notulatus
  • Distichus mahratta
  • Distichus picicornis
  • Distichus puncticollis
  • Distichus uncinatus
  • Dolichoctis chitra
  • Dolichoctis goniodera
  • Dolichoctis marginifer
  • Dolichoctis strita - ssp. striata
  • Dolichoctis vitticollis
  • Dromius orthogonioides
  • Drypta lineola - ssp. lineola
  • Drypta mastersii
  • Dyschirius bengalensis
  • Dyschirius mahratta
  • Dyschirius ordinatus
  • Dyschirius paucipunctus
  • Elaphropus amabilis - ssp. orantus
  • Elaphropus amplians
  • Elaphropus arcuatus
  • Elaphropus ceylanicus
  • Elaphropus charis
  • Elaphropus decoratus
  • Elaphropus diabrachys
  • Elaphropus eueides
  • Elaphropus finitimus
  • Elaphropus granarius
  • Elaphropus horni
  • Elaphropus interpunctatus
  • Elaphropus klugii
  • Elaphropus latus
  • Elaphropus nalandae
  • Elaphropus nilgiricus
  • Elaphropus notaphoides
  • Elaphropus occultus
  • Elaphropus ovatus
  • Elaphropus peryphinus
  • Elaphropus politus - ssp. politus
  • Elaphropus rubescens
  • Elaphropus suturalis
  • Endynomena pradieri
  • Eucolliuris fuscipennis
  • Euplynes marginatus
  • Euschizomerus denticollis
  • Eustra ceylanica
  • Gnaphon costatus
  • Gnathaphanus vulneripennis
  • Gnopheroides pearsoni
  • Harpaglossus opacus
  • Helluodes taprobanae
  • Heteropaussus taprobanensis
  • Holcocoleus melanopus
  • Holcoderus fissus
  • Holcoderus praemorsus
  • Hololeius ceylanicus
  • Hypaetha biramosa - sp. biramosa
  • Hypaetha quadrilineata
  • Hyphaereon consors
  • Hyphaereon hornianus
  • Hyphaereon maculatus
  • Hyphaereon vittatus
  • Idiomorphus guerini
  • Jansenia cirrhidia
  • Jansenia corticata
  • Jansenia laeticolor
  • Jansenia stellata
  • Jansenia westermanni
  • Lachnothorax biguttatus
  • Lasiocera coromandelica
  • Lebia dichroma
  • Lebia exsanguis
  • Lebia leucaspis
  • Lebia lunigera
  • Lebia monostigma
  • Leleuporella sexangulata
  • Lionychus albivittis
  • Lionychus horni
  • Lophyra cancellata
  • Lophyra catena - ssp. catena, insularis
  • Lorostema alutacea - ssp. alutacea, spinipennis
  • Loxocrepis ruficeps
  • Loxoncus discophorus
  • Loxoncus microgonus
  • Loxoncus nagpurensis
  • Loxoncus renitens
  • Loxoncus schmidti
  • Macrocheilus bensoni
  • Macrocheilus niger
  • Masoreus orientalis
  • Mastax euanthes
  • Mastax histrio
  • Melaenus piger
  • Melanospilus andrewesi
  • Metacolpodes buchannani
  • Metazuphium spinangulis
  • Microlestes demessus - ssp. demessus
  • Microlestes inconspicuus
  • Microlestes xanthopus
  • Mimocolliuris pilifera
  • Mimocolliuris stigma
  • Miscelus javanus
  • Miscelus unicolor
  • Mochtherus tetraspilotus
  • Morion cucujoides
  • Morion orientalis
  • Myriochila distinguenda
  • Myriochila fastidiosa - ssp. fastidiosa
  • Myriochila undulata
  • Nanodiodes piceus
  • Nanodiodes westermanni
  • Neocollyris aenea
  • Neocollyris ceylonica
  • Neocollyris crassicornis - ssp. crassicornis
  • Neocollyris planifrontoides
  • Neocollyris plicaticollis
  • Neocollyris punctatella
  • Neocollyris saundersii - ssp. laetior, saundersii
  • Neocollyris sedlaceki
  • Neocollyris tuberculata
  • Neocollyris vedda
  • Nototachys comptus
  • Omphra hirta
  • Omphra pilosa
  • Omphra rufipes
  • Oodes angustatus
  • Oodes bivittatus
  • Oodes taprobanae
  • Oodes xanthochilus
  • Ooidius advolans
  • Oosoma gyllenhalii
  • Oosoma semivittatum
  • Ophionea ceylonica
  • Ophionea indica
  • Ophionea interstitialis
  • Ophionea nigrofasciata - ssp. nigrofasciata
  • Ophoniscus insulicola
  • Ophoniscus iridulus
  • Orthogonius acutangulus
  • Orthogonius batesi
  • Orthogonius femoralis
  • Orthogonius fugax
  • Orthogonius longicornis
  • Orthogonius ovatulus
  • Orthogonius parallelus
  • Orthogonius planiger
  • Orthogonius schaumi
  • Orthogonius srilankaicus
  • Oxylobus lateralis - ssp. designans
  • Oxylobus ovalipennis
  • Oxylobus porcatus
  • Paradromius steno
  • Paraphaea binotata
  • Parena nigrolineata
  • Parophonus compositus
  • Parophonus cyaneotinctus
  • Parophonus javanus
  • Parophonus lividus
  • Parophonus nagpurensis - ssp. curvatus
  • Paussus desneuxi
  • Paussus escherichi
  • Paussus fletcheri
  • Paussus horni
  • Paussus pacificus
  • Paussus politus
  • Peliocypas catenatus
  • Peliocypas euproctoides
  • Peliocypas fuscus
  • Peliocypas intermedius
  • Peliocypas levipennis
  • Peliocypas malleus
  • Peliocypas oryctus
  • Peliocypas repandus
  • Peliocypas trigonus
  • Pelocharis remyi
  • Pentagonica ceylonica
  • Pentagonica erichsoni
  • Pentagonica horni
  • Pentagonica pallipes
  • Pentagonica ruficeps
  • Pentagonica ruficollis
  • Pentagonica venusta
  • Perigona castanea
  • Perigona nigriceps
  • Perigona nigricollis
  • Perigona nigrifrons
  • Perigona plagiata
  • Perigona sinuaticollis
  • Perigona tronqueti
  • Perileptus ceylanicus
  • Perileptus indicus
  • Peripristus ater
  • Pheropsophus bimaculatus
  • Pheropsophus catoirei
  • Pheropsophus chaudoiri
  • Pheropsophus discicollis
  • Pheropsophus lissoderus
  • Pheropsophus occipitalis
  • Pheropsophus picicollis
  • Physocrotaphus ceylonicus
  • Physodera eschscholtzii
  • Planetes bimaculatus
  • Planetes elegans
  • Planetes ruficeps
  • Planetes ruficollis
  • Planetes simplex
  • Platymetopus flavilabris
  • Platymetopus keiseri
  • Platymetopus pictus
  • Platymetopus rugosus
  • Platytarus boysii
  • Prothyma proxima
  • Protocollyris planifrons
  • Pseudoclivina memnonia
  • Pseudognathaphanus dispellens
  • Pseudognathaphanus punctilabris
  • Pseudognathaphanus rusticus
  • Pterostichus atratus
  • Scarites ceylonicus
  • Scarites indus
  • Scarites selene
  • Selina westermanni
  • Sericoda ceylonica
  • Siagona depressa - ssp. depressa
  • Siagona fabricii
  • Siagona plana
  • Somotrichus unifasciatus
  • Stenolophus opaculus
  • Stenolophus pallipes
  • Stenolophus polygenus
  • Stenolophus quinquepustulatus
  • Stenolophus rectifrons
  • Stenolophus smaragdulus
  • Stomonaxellus ceylanensis
  • Styphlomerus fusciceps
  • Syleter porphyreus
  • Syleter validus
  • Syntomus quadripunctatus
  • Tachys brachys
  • Tachys euryodes
  • Tachys fasciatus - ssp. faciatus
  • Tachys impressipennis
  • Tachys incertus
  • Tachys opalescens
  • Tachys quadrillum
  • Tachys sexguttatus
  • Tachys tropicus
  • Tachys truncatus
  • Tachys umbrosa
  • Tantillus brunneus
  • Tantillus vittatus
  • Tetragonoderus cursor
  • Tetragonoderus fimbriatus
  • Tetragonoderus notaphioides
  • Tetragonoderus quadrinotatus
  • Trichotichnus hiekei
  • Trichotichnus lamprus
  • Trichotichnus lucens
  • Trichotichnus marginalis
  • Trichotichnus nitens
  • Trichotichnus pseudolucens
  • Tricondyla coriacea
  • Tricondyla femorata
  • Tricondyla granulifera
  • Tricondyla nigripalpis
  • Trigonotoma indica
  • Trilophidius impunctatus
  • Trilophus arcuatus
  • Xenodochus mediocris
  • Zuphium dabreui
  • Zuphium erebeum
  • Zuphium olens

Family: Cerambycidae - Longhorn beetles

Family: Cerylonidae - Minute bark beetles

Family: Chrysomelidae - Leaf beetles

  • Amblispa dohrnii
  • Aspidimorpha dorsata
  • Aspidimorpha furcata
  • Bruchidius anderssoni
  • Bruchidius brincki
  • Bruchidius cingalicus
  • Bruchidius mendosus
  • Bruchidius minutissimus
  • Bruchidius nalandus
  • Bruchidius trifasciatus
  • Bruchidius walkeri
  • Callosobruchus bicalcaratus
  • Callosobruchus chinensis
  • Callosobruchus gibbicollis
  • Callosobruchus nigripennis
  • Caryedon serratus
  • Cassida ceylonica
  • Cassida circumdata
  • Cassida dorsonotata
  • Cassida imbecilla
  • Cassida obtusata
  • Chiridopsis marginata
  • Eutornus jansoni
  • Galerucella placida
  • Medythia suturalis
  • Monolepta cognata
  • Monolepta descripta
  • Monolepta juno
  • Monolepta oculata
  • Monolepta sexlineata
  • Platycorynus peregrinus
  • Sagra femorata
  • Spermophagus aeneipennis
  • Spermophagus albosparsus
  • Spermophagus cederholmi
  • Spermophagus mannarensis
  • Spermophagus niger
  • Spermophagus pfaffenbergeri

Family: Cleridae - Checkered beetles

Family: Coccinellidae - Lady beetles

Family: Cucujidae - Flat bark beetles

  • Cryptolestes ferrugineus
  • Cryptolestes klapperichi
  • Laemophloeus incertus
  • Narthecius bicolor
  • Notolaemus lewisi
  • Placonotus orientalis
  • Placonotus subtestaceus
  • Placonotus torsus
  • Xylolestes krombeini

Family: Curculionidae - Snout and Bark bBeetles

  • Acacacis zeylanicus
  • Acicnemis dorsonotata
  • Acicnemis horni
  • Acicnemis mansueta
  • Acicnemis reversa
  • Aclees birmanus
  • Alcidodes comptus
  • Alcidodes erosus
  • Alcidodes fornicatus
  • Alcidodes lugubris
  • Alcidodes magnificus
  • Alcidodes notabilis
  • Alcidodes texatus
  • Alcidodes virgatus
  • Ambrosiodmus asperatus
  • Amphialus agrestis
  • Amphialus communicans
  • Amphialus descriminans
  • Amphialus turgidus
  • Amphorygma ceylonensis
  • Arixyleborus granulifer
  • Arixyleborus rugosipes
  • Astycus aequalis
  • Astycus apicatus
  • Astycus armatipes
  • Astycus bilineatus
  • Astycus canus
  • Astycus cinereus
  • Astycus cinnamomeus
  • Astycus horni
  • Astycus immunis
  • Astycus lewisi
  • Astycus suturalis
  • Atinella senex
  • Balaninus c-album
  • Blosyrus inaequalis
  • Calandra longicollis
  • Calandra sordidus
  • Camptorrhinus reversa
  • Cleonus arenarius
  • Cleonus gyllenhali
  • Cleonus ophinotus
  • Cleonus zebra
  • Cnestus magnus
  • Coccotrypes advena
  • Coccotrypes cyperi
  • Coccotrypes rugicollis
  • Coccotrypes theae
  • Coccotrypes variabilis
  • Coccotrypes vulgaris
  • Cosmopolites sordidus
  • Cossonus ambiguus
  • Cossonus disciferus
  • Cossonus hebes
  • Cossonus horni
  • Craniodicticus mucronatus
  • Cratopus sinhalensis
  • Crossotarsus minor
  • Crossotarsus saundersi
  • Cryphalogenes euphorbiae
  • Cryphalogenes exiguus
  • Cryphalomorphus opacus
  • Cryphalops capucinus
  • Cryphalops quadridens
  • Cryphalus fuliginosus
  • Cryphalus neglectus
  • Cryphalus nigricans
  • Cryphalus palawanus
  • Cryphalus subvestitus
  • Cryphalus tetricus
  • Cryphalus vestitus
  • Cryptorhynchus assimilans
  • Cryptorhynchus declaratus
  • Curculio arenarius
  • Curculio c-album
  • Curculio curvicornis
  • Curculio dentifer
  • Curculio lineolatus
  • Curculio lugubris
  • Curculio motschulskyi
  • Curculio rufimanus
  • Curculio transmarinus
  • Curculio viridanus
  • Cyrtotrachelus rufopectinipes
  • Dacryophthorus brincki
  • Dendroctonomorphus muricatus
  • Dereodus prasinus
  • Dereodus sparsus
  • Desmidophorinus fasciculicollis
  • Diamerus curvifer
  • Diocalandra subfasciatus
  • Dryocoetes flavicornis
  • Dryocoetes taprobanus
  • Dryophthorus setulosus
  • Eccoptopterus spinosus
  • Epicalus virgatus
  • Episomus figuratus
  • Episomus fimbriatus
  • Episomus pyriformis
  • Euwallacea bicolor
  • Euwallacea dilatatiformis
  • Euwallacea interjectus
  • Euwallacea piceus
  • Exilis horni
  • Ficicis despectus
  • Genyocerus albipennis
  • Hylurgus concinnulus
  • Hypoborus cinereotestaceus
  • Hypocryphalus cylindripennis
  • Hypocryphalus glabratus
  • Hypocryphalus interponens
  • Hypocryphalus mangiferae
  • Hypothenemus areccae
  • Hypothenemus bicinctus
  • Hypothenemus eruditus
  • Hypothenemus externedentatus
  • Hypothenemus seriatus
  • Ischnopus taprobanus
  • Liparthrum brincki
  • Macrorhyncolus crassitarsis
  • Mechistocerus assimilans
  • Mechistocerus declaratus
  • Megarrhinus cingalensis
  • Mesites subvittatus
  • Mesites suturalis
  • Myllocerus angulatipes
  • Myllocerus canescens
  • Myllocerus curvicornis
  • Myllocerus delicatulus
  • Myllocerus dentifer
  • Myllocerus fringilla
  • Myllocerus retratiens
  • Myllocerus spurcatus
  • Myllocerus subfasciatus
  • Myllocerus transmarinus
  • Myllocerus undatus
  • Myllocerus variegatus
  • Myllocerus viridanus
  • Myllocerus zeylanicus
  • Myocalandra exarata
  • Nassophasis foveata
  • Nassophasis pictipes
  • Neodiamerus granulicollis
  • Niphades vexatus
  • Odoiporus longicollis
  • Odontobarus hodiernus
  • Olonthogaster regalis
  • Omotemnus introducens
  • Omphasus nalandae
  • Orthosinus salutarius
  • Orthosinus sculpticollis
  • Orthosinus velatus
  • Otidognathus meridionalis
  • Oxydema fusiforme
  • Peltotrachelus ovis
  • Pentarthrum wollastoni
  • Phaenomerus sundevalli
  • Phloeoditica curtus
  • Phloeophagosoma atratum
  • Phloeophagosoma morio
  • Phloeophagus cossonoides
  • Phloeosinus detersus
  • Phloeotrogus obliquecaudata
  • Phrygena ephippiata
  • Piazomias aequalis
  • Piazomias immunis
  • Platypus furcatus
  • Platypus latifinis
  • Platypus rotundicauda
  • Platypus solidus
  • Platyrrhinus marmoratus
  • Platytrachelus ovis
  • Polytus mellerborgi
  • Prodioctes haematicus
  • Prodioctes singhalensis
  • Psilosomus hebes
  • Ptilopodius ceylonicus
  • Ptochus imbricatus
  • Ptochus limbatus
  • Ptochus planoculis
  • Ptochus pyriformis
  • Rhabdocnemis maculata
  • Rhinoncus paganus
  • Rhynchaenus c-album
  • Rhynchophorus ferrugineus
  • Rhynchophorus introducens
  • Rhyncolus ater
  • Rhyncolus cossonoides
  • Rhyncolus linearis
  • Rhyncolus taciturnus
  • Scolytomimus dilutus
  • Scolytomimus mimusopis
  • Scolytomimus rectus
  • Sipalinus gigas
  • Sipalus hypocrita
  • Sipalus porosus
  • Sitophilus exarata
  • Sitophilus mellerborgi
  • Sitophilus subfasciatus
  • Sphaerotrypes cristatus
  • Sphaerotrypes ranasinghei
  • Sphenophorus maculata
  • Sphenophorus planipennis
  • Sphenophorus sordidus
  • Stereotribus incisus
  • Stereotribus tuberculifrons
  • Strattis biguttatus
  • Strattis vestigialis
  • Strophosomus suturalis
  • Sueus niisimai
  • Sympiezomias kraatzi
  • Tanymecus curviscapus
  • Treptoplatypus solidus
  • Trigonocolus cingalensis
  • Trochorhopalus discifer
  • Trochorhopalus leucogrammus
  • Tylodes semicollis
  • Webbia ceylonae
  • Xyleborinus andrewesi
  • Xyleborinus exiguus
  • Xyleborus affinis
  • Xyleborus arquatus
  • Xyleborus comptus
  • Xyleborus costipennis
  • Xyleborus cristatuloides
  • Xyleborus cuneolosus
  • Xyleborus dilatatiformis
  • Xyleborus doliaris
  • Xyleborus emarginatus
  • Xyleborus figuratus
  • Xyleborus fornicatior
  • Xyleborus fornicates
  • Xyleborus haberkorni
  • Xyleborus insitivus
  • Xyleborus perforans
  • Xyleborus pseudocitri
  • Xyleborus rimulosus
  • Xyleborus seminitens
  • Xyleborus similis
  • Xyleborus tenella
  • Xylosandrus compactus
  • Xylosandrus crassiusculus
  • Xylosandrus discolor
  • Xylosandrus pygmaeus

Family: Dryopidae - Long-toed water beetles

Family: Dytiscidae - Predaceous diving beetles

Family: Elateridae - Click beetles

  • Anchastus humeralis
  • Calais speciosus
  • Ceropectus cederholmi
  • Conoderus pruinosus
  • Cryptalaus sordidus
  • Dicronychus stolatus
  • Drasterius collaris
  • Elius dilatatus
  • Lanelater fuscipes
  • Lanelater sobrinus
  • Melanotus vermiculatus
  • Meristhus ceylonensis
  • Mulsanteus hirtellus
  • Negastrius anderssoni
  • Negastrius brincki
  • Paracardiophorus fuscipennis
  • Rismethus diodesmoides
  • Rismethus minusculus

Family: Elmidae - Riffle beetles

  • Ancyronyx jaechi
  • Cephalolimnius ater
  • Graphelmis ceylonica
  • Ilamelmis brunnescens
  • Ilamelmis crassa
  • Ilamelmis foveicollis
  • Ilamelmis starmuhlneri
  • Ilamelmis cederholmi
  • Ordobrevia flavolineata
  • Ordobrevia fletcheri
  • Podelmis aenea
  • Podelmis ater
  • Podelmis metallica
  • Podelmis ovalis
  • Podelmis quadriplagiata
  • Podelmis viridiaenea
  • Potamophilinus costatus
  • Potamophilinus impressicollis
  • Stenelmis anderssoni
  • Stenelmis brincki
  • Negastrius anderssoni
  • Negastrius brincki
  • Taprobanelmis carinata
  • Zaitzeviaria bicolor
  • Zaitzeviaria zeylanica

Family: Endomychidae - Handsome fungus beetles

  • Ancylopus ceylonicus
  • Chondria minima
  • Endocoelus orbicularis
  • Eumorphus quadriguttatus - ssp. pulchripes
  • Idiophyes ceylonica
  • Mycetina castanea
  • Ohtaius lunulatus
  • Saula nigripes
  • Stenotarsus tomentosus
  • Trochoideus desjardinsi

Family: Erotylidae - Pleasing fungus beetles

Family: Geotrupidae - Earth-boring dung beetles

Family: Gyrinidae - Whirligig beetles

  • Aulonogyrus obliquus
  • Gyrinus convexiusculus
  • Gyrinus sericeolimbatus
  • Orectochilus dilatatus
  • Orectochilus discifer
  • Orectochilus fraternus
  • Orectochilus indicus
  • Orectochilus wehnckei
  • Porrorhynchus brevirostris

Family: Histeridae - Clown beetles

  • Acritus copricola
  • Acritus tuberisternus
  • Atholus coelestis
  • Atholus daldorffi
  • Bacanius ambiguus
  • Chaetabraeus granosus
  • Chaetabraeus orientalis
  • Chaetabraeus paria
  • Chalcionellus pulchellus
  • Cypturus aenescens
  • Epiechinus taprobanae
  • Halacritus alutiger
  • Hister javanicus
  • Hister trigonifrons
  • Hololepta elongata
  • Liopygus minutus
  • Liopygus subsuturalis
  • Nasaltus chinensis
  • Nasaltus orientalis
  • Pachylister lutarius
  • Pachylister scaevola
  • Platylister desinens
  • Platylomalus biarculus
  • Platylomalus oceanitis
  • Tribalus colombius
  • Trypeticus bombacis

Family: Hybosoridae - Scavenger scarab beetles

  • Besuchetostes besucheti
  • Besuchetostes loebli
  • Besuchetostes mussardi
  • Besuchetostes peradeniyae
  • Besuchetostes taprobanae
  • Hybosorus orientalis
  • Phaeochrous compactus
  • Phaeochrous elevatus
  • Phaeochrous emarginatus
  • Pterorthochaetes haroldi

Family: Hydrophilidae - Water scavenger beetles

  • Berosus pulchellus
  • Berosus undatus
  • Cercyon connivens
  • Cercyon lineolatus
  • Cercyon nigriceps
  • Cercyon subsolanus
  • Coelostoma stultum
  • Dactylosternum abdominale
  • Enochrus esuriens
  • Helochares mundus
  • Hydrophilus bilineatus - ssp. caschmirensis
  • Paroosternum sorex
  • Pelthydrus jengi
  • Regimbartia attenuata
  • Sphaeridium quinquemaculatum
  • Sternolophus inconspicuus
  • Sternolophus rufipes

Family: Lampyridae - Fireflies

Family: Lucanidae - Stag beetles

  • Aegus cehlifer - ssp. kandiensis
  • Aegus dilatus
  • Calcodes carinatus
  • Dinonigidius ahenobarbus
  • Figulus aratus
  • Figulus horni
  • Figulus howei
  • Figulus interruptus
  • Neolucanus sinicus - ssp. championi
  • Platyfigulus scorpio
  • Prosopocoilus henryi

Family: Lycidae - Net-winged beetles

Family: Lymexylidae - Ship-timber beetles

Family: Meloidae - Blister beetles

Family: Melolonthidae - White grubs

  • Apogonia aequabilis
  • Apogonia cava
  • Apogonia comosa
  • Apogonia coriacea
  • Apogonia feruginea
  • Apogonia fulvosetosa
  • Apogonia gracilis
  • Apogonia laevissima
  • Apogonia lateralis
  • Apogonia lurida
  • Apogonia nana
  • Apogonia nietneri
  • Apogonia prolixa
  • Apogonia rauca
  • Apogonia solida
  • Autoserica dubia
  • Autoserica pubescens
  • Autoserica srilanka
  • Brahmina flavipennis
  • Dichelomorpha glabrilinea
  • Dichelomorpha punctuligera
  • Engertia maculosa
  • Holotrichia ceylonensis
  • Holotrichia convexifrons
  • Holotrichia danielssoni
  • Holotrichia disparilis
  • Holotrichia eurystoma
  • Holotrichia furcifer
  • Holotrichia inducta
  • Holotrichia kandulawai
  • Holotrichia opuana
  • Holotrichia parva
  • Holotrichia remorata
  • Holotrichia reynaudi
  • Holotrichia rufoflava
  • Holotrichia schmitzi
  • Holotrichia semitomentosa
  • Holotrichia serrata
  • Holotrichia setosa
  • Idiochelyna pectoralis
  • Lepidiota ferruginosa
  • Lepidiota horni
  • Lepidiota pinguis
  • Maladera breviata
  • Maladera calcarata
  • Maladera cardoni
  • Maladera chalybaea
  • Maladera cinnaberina
  • Maladera coxalis
  • Maladera fistulosa
  • Maladera immunita
  • Maladera implicata
  • Maladera insanabilis
  • Maladera rotunda
  • Maladera rufocuprea
  • Maladera setosa
  • Maladera singhalensis
  • Maladera straba
  • Maladera weligamana
  • Maladera pinguis
  • Neoserica bombycina
  • Neoserica sexfoliata
  • Neoserica splendifica
  • Periserica fracta
  • Periserica nigripennis
  • Periserica picta
  • Rhizotrogus aequalis
  • Rhizotrogus hirtipectus
  • Rhizotrogus sulcifer
  • Schizonycha ruficollis
  • Schizonycha singhalensis
  • Schoenherria maculosa
  • Selmaladera nitida
  • Selmaladera pusilla
  • Selmaladera scutellaris
  • Selmaladera sericea
  • Serica distincticornis
  • Serica fusa
  • Serica hamifera
  • Serica interrupta
  • Serica lurida
  • Serica maculicauda
  • Serica maculifera
  • Serica nana
  • Serica nepalensis
  • Serica rubescens
  • Serica semicincta
  • Serica ventriosa
  • Sophrops costatus
  • Sophrops eurystomus
  • Sophrops singhalensis
  • Stephanopholis cribricollis
  • Stephanopholis ruficundus
  • Stephanopholis singalensis

Family: Nitidulidae - Sap beetles

Family: Passalidae - Bess beetles

  • Aceraius comptoni
  • Aceraius meyeri
  • Episphenus comptoni
  • Episphenus flachi
  • Episphenus moorei
  • Leptaulax bicolor
  • Macrolinus batesi
  • Macrolinus crenatipennis
  • Macrolinus obesus
  • Macrolinus rotundifrons
  • Macrolinus waterhousei
  • Stephanocephalus redtenbacheri
  • Taeniocerus bicanthatus

Family: Ptiliidae - Feather-wing beetles

Family: Ptinidae - Spider beetles

Family: Rhynchitidae - Tooth-nosed snout weevils

  • Auletobius ceylonicus
  • Auletobius maculatus
  • Auletobius subgranulatus
  • Auletobius testaceipennis
  • Deporaus ceylonensis
  • Deporaus marginatus
  • Deporaus rufipallens
  • Eugnamptus marginatus
  • Rhynchites amictus
  • Rhynchites ceylonensis
  • Rhynchites clavatus
  • Rhynchites fasciatus
  • Rhynchites flavipedestris
  • Rhynchites flavipes
  • Rhynchites nalandaicus
  • Rhynchites parvulus
  • Rhynchites restituens
  • Rhynchites suffundens
  • Rhynchites vestitus

Family: Rutelidae - Flower beetles

  • Adoretus bicaudatus
  • Adoretus celogaster
  • Adoretus corpulentus
  • Adoretus duplicatus
  • Adoretus erythrocephalus
  • Adoretus feminalis
  • Adoretus leo
  • Adoretus sorex
  • Adoretus suturalis
  • Adoretus versutus
  • Adoretus chloronota
  • Adoretus conformis
  • Adoretus discalis
  • Adoretus discors
  • Adoretus dussumieri
  • Adoretus fracta
  • Adoretus gravida
  • Adoretus illusa
  • Adoretus infantilis
  • Adoretus pellucida
  • Adoretus procrastinator
  • Adoretus ruficapilla
  • Adoretus superflua
  • Adoretus varicolor
  • Adoretus viridis
  • Adoretus walkeri
  • Callistethus bugnioni
  • Callistethus chloromelus
  • Callistethus princeps
  • Lepadoretus compressus
  • Lepadoretus ermineus
  • Lepadoretus mavis
  • Lepadoretus nietneri
  • Micranomala cingalensis
  • Mimela macleayana
  • Mimela mundissima
  • Parastasia basalis
  • Parastasia cingala
  • Parastasia lobata
  • Popillia discalis
  • Rhamphadoretus sorex
  • Rhinyptia gilleti
  • Rhinyptia meridionalis
  • Singhala hindu
  • Singhala tenella
  • Trogonostomus ursus

Family: Salpingidae - Narrow-waisted bark beetles

Family: Scarabaeidae - Scarab beetles

  • Agestrata orichalca - ssp. nigrita
  • Alissonotum piceum
  • Anthracophora crucifera
  • Blabephorus pinguis
  • Caccobius aterrimus
  • Caccobius diminutivus
  • Caccobius indicus
  • Caccobius meridionalis
  • Caccobius rufipennis
  • Caccobius unicornis
  • Caccobius vulcanus
  • Campsiura nigripennis - ssp. cingalensis
  • Catharsius capucinus
  • Catharsius granulatus
  • Catharsius molossus
  • Catharsius pithecius
  • Chaetadoretus infans
  • Chaetadoretus lasiopygyus
  • Chaetadoretus mus
  • Chaetadoretus rugosus
  • Chaetadoretus silonicus
  • Chaetadoretus singhalensis
  • Chiron cylindrus
  • Cleptocaccobius durantoni
  • Cleptocaccobius inermis
  • Clinteria ceylonensis
  • Clinteria chloronota
  • Clinteria coerulea - ssp. megaspilota
  • Clinteria imperialis
  • Clinteria jansoni
  • Clinteria keiseri
  • Clinteria klugi - ssp. rufipennis
  • Clinteria pantherina
  • Clyster itys
  • Coenochilus taprobanicus
  • Copris fricator
  • Copris repertus
  • Copris signatus
  • Copris sodalis
  • Dasyvalgus addendus
  • Delopleurus parvus
  • Dicheros bimacula
  • Digitonthophagus bonasus
  • Digitonthophagus gazella
  • Dipelicus bidens
  • Dipelicus daedalus
  • Dipelicus hircus
  • Eophileurus cingalensis - ssp. decatenatus
  • Euselates scenica
  • Gametis versicolor
  • Garreta smaragdifer
  • Glycosia tricolor
  • Glycyphana horsfieldi - ssp. aurulenta
  • Gymnopleurus cyaneus
  • Gymnopleurus gemmatus
  • Gymnopleurus koenigi
  • Gymnopleurus miliaris
  • Gymnopleurus parvus
  • Gymnopleurus sericeifrons
  • Haroldius herrenorum
  • Haroldius krali
  • Heliocopris bucephalus
  • Heteronychus krombeini
  • Heterorrhina elegans
  • Heterorrhina planat
  • Kheper erichsoni
  • Kheper sanctus
  • Liatongus rhadamistus
  • Neosisyphus tarantula
  • Ochicanthon ceylonicus
  • Ochicanthon cingalensis
  • Onitis crenatus
  • Onitis philemon
  • Onitis singhalensis
  • Onitis subopacus
  • Onthophagus capitosus
  • Onthophagus centricornis
  • Onthophagus cervus
  • Onthophagus cryptogenus
  • Onthophagus dama
  • Onthophagus difficilis
  • Onthophagus falsus
  • Onthophagus favrei
  • Onthophagus (Palaeonthophagus) fissicornis
  • Onthophagus frontalis
  • Onthophagus fuscopunctatus
  • Onthophagus gemma
  • Onthophagus gravis
  • Onthophagus hastifer
  • Onthophagus heterorrhinus
  • Onthophagus hystrix
  • Onthophagus keiseri
  • Onthophagus laevigatus
  • Onthophagus ludio
  • Onthophagus luridipennis
  • Onthophagus martialis
  • Onthophagus militaris
  • Onthophagus modestus
  • Onthophagus negligens
  • Onthophagus occipitalis
  • Onthophagus ochreatus
  • Onthophagus oculatus
  • Onthophagus parvulus
  • Onthophagus politus
  • Onthophagus pusillus
  • Onthophagus pygmaeus
  • Onthophagus (Colobonthophagus) quadridentatus
  • Onthophagus (Serrophorus) rectecornutus
  • Onthophagus rufulgens
  • Onthophagus regalis
  • Onthophagus solmani
  • Onthophagus sparsepunctatus
  • Onthophagus spinifex
  • Onthophagus taprobanus
  • Onthophagus tritinctus
  • Onthophagus turbatus
  • Onthophagus unifasciatus
  • Onthophagus vacca
  • Oreoderus insularis
  • Orphnus parvus
  • Oryctes gnu
  • Oryctes monoceros
  • Oryctes rhinoceros
  • Panelus ceylonicus
  • Panelus fallax
  • Panelus imitator
  • Panelus pernitidus
  • Panelus puncticollis
  • Panelus setosus
  • Paragymnopleurus melanarius
  • Phaedotrogus ceylonicus
  • Phalops divisus
  • Phyllognathus dionysius
  • Proagoderus pactolus
  • Protaetia alboguttata
  • Protaetia aurichalcea
  • Protaetia ceylanica
  • Protaetia cupripes - ssp. germar
  • Panelus horni
  • Panelus rufocuprea
  • Panelus whitehousei
  • Panelus wiebesi
  • Scarabaeus gangeticus
  • Scarabaeus sanctus
  • Sisyphus crispatus - ssp. hirtus
  • Sisyphus indicus
  • Sisyphus longipes
  • Spilophorus cretosus
  • Taeniodera flavomaculata
  • Taeniodera halyi
  • Taeniodera quadrivittata
  • Thaumastopeus ceylonicus
  • Tibiodrepanus hircus
  • Tibiodrepanus setosus
  • Xylotrupes gideon - ssp. socrates
  • Xylotrupes meridionalis
  • Xylotrupes taprobanus

Family: Scydmaenidae - Ant-like stone beetles

Family: Silvanidae - Silvan flat bark beetles

Family: Staphylinidae - Rove beetles

  • Anotylus exiguus
  • Anotylus glareosus
  • Anotylus hybridus
  • Anotylus insecatus
  • Anotylus latiusculus
  • Anotylus tetracarinatus
  • Baeocera longicornis
  • Batribolbus palpator
  • Batrisiella satoi
  • Carpelimus fuliginosus
  • Cilea limbifer
  • Erymus gracilis
  • Oligota parva
  • Oxytelopsis pseudopsina
  • Oxytelus bengalensis
  • Oxytelus incisus
  • Oxytelus migrator
  • Oxytelus nigriceps
  • Oxytelus puncticeps
  • Oxytelus varipennis
  • Scaphisoma stictum
  • Scopaeus testaceus
  • Stenus bicornis
  • Stenus cordatus
  • Stenus melanarius
  • Stenus serupeus
  • Stenus solutus

Family: Tenebrionidae - Darkling beetles

  • Alphitobius laevigatus
  • Amarygmus picicornis
  • Ceropria induta
  • Corticeus cephalotes
  • Eutochia aptera
  • Falsandrosus tetrops
  • Gonocephalum depressum
  • Laena formaneki
  • Leichenum canaliculatum
  • Pimelia undulata
  • Platydemoides brincki
  • Tribolium castaneum
  • Uloma bituberosa
  • Uloma polita
  • Ulomina carinata

Family: Trogidae - Hide beetles

Family: Vesperidae - Vesperid longicorn beetles

Family: Zopheridae - Ironclad beetles

  • Bitoma siviana
  • Cebia rugosa
  • Microprius opacus
  • Monomma brunneum
  • Nematidium elongatum
  • Trachypholis hispidum

Notes

    References

    1. Polilov, A.A (March 2008). "Anatomy of the smallest coleoptera, featherwing beetles of the tribe nanosellini (Coleoptera, Ptiliidae), and limits of insect miniaturization". Entomological Review , Volume 88, Issue 1. pp. pp 26–33. Retrieved 22 September 2016.
    2. Mannakkara, Amani., Makihara, K. "Checklist of longicorn coleoptera of Sri Lanka (1) Vesperidae and Cerambycidae excluding Lamiinae" (PDF). ffpri. Retrieved 22 September 2016.
    3. "Scirtidae (Coleoptera), aquatic larvae, Sri Lanka". Pest net. Retrieved 22 September 2016.
    4. Wellappuli Arachchi, Supun. "Common Insects Of Sri Lanka" (PDF). Butterfly Conservation Society of Sri Lanka. Retrieved 22 September 2016.
    5. "STUDY ON COLEOPTERAN FAUNA IN SOIL AND LITTER OF WAGA NATURAL. FOREST RESERVE IN SRI LANKA" (PDF). Open University of Sri Lanka. Retrieved 22 September 2016.
    6. "Predatory Coccinellids (Coleoptera: Cocinellidae) of Vegetables in Sri Lanka" (PDF). Tropical Agricultural Research Vol.19. pp. 69–77. Retrieved 22 September 2016.
    7. "Rhynchophorus ferrugineus attractants" (PDF). National Science Foundation of Sri Lanka. Retrieved 22 September 2016.
    8. Bambaradeniya, Channa N. B. "The Fauna of Sri Lanka: Status of Taxonomy, Research, and Conservation". Amazon.com. Google books. Retrieved 23 January 2016.
    9. "Coleoptera in Sri Lanka". insectoid. Retrieved 22 September 2016.
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